What is the most effective revision to make in sentence 23?

(23) For example, the iguana, a WebMD article states, sneezes more often than any other animal and more efficiently.

For example, the iguana sneezes more often and more efficiently than any other animal is what a WebMD article states.

For example, a WebMD article states that the iguana sneezes more often and more efficiently than any other animal.

For example, a WebMD article states this, the iguana sneezes more often and more efficiently than any other animal.

No revision is needed in this sentence.